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Figure 6.1: Coat color in animals is an example of a trait that affected by more than one locus.

The principles of genetic analysis that we have described for a single locus can be extended to the study of two genes simultaneously.   Analysis of two loci in parallel is required for genetic mapping and can also reveal gene interactions.  These techniques can be very useful for both basic and applied research. Before discussing these techniques, we will first revisit Mendel and his groundbreaking experiments.